| WangHeng-Chang等也表明，染色体片段和落后染色体可以导致微核形成。实验中末期Ⅰ，末期Ⅱ及四分体时期的微核来源于分裂过程中染色体片段和落后染色体。观察还发现落后染色体发生的频率随着减数分裂进行逐渐降低，这可能与进入末期，染色体片段和落后染色体随机分布或丢失有关，与此同时，落后染色体的移动速度较慢，可能是导致减数分裂不同步的主要原因。
四分体时期，出现染色体多极分裂等异常现象。根据李雅轩、王祖秀、唐仙英、胡丹艳等的观点，作者推断龙眼染色体多极分裂，形成多分孢子，可能与纺锤体定位异常有关。但本实验中未观察到清晰的纺锤体形态。有关纺锤体对减数分裂染色体异常行为的作用机制，有待更深入研究。
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